Edgar Filing: APPLERA CORP

Ownership Submission

FORM 4

Check this box if no longer subject to Section 16. Form 4 or Form 5 obligations may continue. See Instruction 1(b).

UNITED STATES SECURITIES AND EXCHANGE COMMISSION
Washington, D.C. 20549
STATEMENT OF CHANGES IN BENEFICIAL OWNERSHIP OF SECURITIES

Filed pursuant to Section 16(a) of the Securities Exchange Act of 1934, Section 17(a) of the Public Utility Holding Company Act of 1935 or Section 30(h) of the Investment Company Act of 1940

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3235-0287

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January 31, 2005

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(Print or Type Responses)

1. Name and Address of Reporting Person ^*

NAYYAR SANDEEP

2. Issuer Name and Ticker or Trading Symbol
APPLERA CORP [ABI/CRA]

5. Relationship of Reporting Person(s) to Issuer
(Check all applicable)

_____ Director	_____ 10% Owner
__X__ Officer (give title below)	_____ Other (specify below)
Assistant Controller

(Last)

(First)

(Middle)

APPLERA CORP - APPLIED BIOSYSTEMS GROUP, 850 LINCOLN CENTRE DRIVE

3. Date of Earliest Transaction (Month/Day/Year)
11/06/2006

(Street)

FOSTER CITY, CA 94404

4. If Amendment, Date Original Filed(Month/Day/Year)

6. Individual or Joint/Group Filing(Check Applicable Line)
_X_ Form filed by One Reporting Person
___ Form filed by More than One Reporting Person

(City)

(State)

(Zip)

Table I - Non-Derivative Securities Acquired, Disposed of, or Beneficially Owned

1.Title of Security (Instr. 3)	2. Transaction Date (Month/Day/Year)	2A. Deemed Execution Date, if any (Month/Day/Year)	3. Transaction Code (Instr. 8)		4. Securities Acquired (A) or Disposed of (D) (Instr. 3, 4 and 5)			5. Amount of Securities Beneficially Owned Following Reported Transaction(s) (Instr. 3 and 4)	6. Ownership Form: Direct (D) or Indirect (I) (Instr. 4)	7. Nature of Indirect Beneficial Ownership (Instr. 4)
1.Title of Security (Instr. 3)	2. Transaction Date (Month/Day/Year)	2A. Deemed Execution Date, if any (Month/Day/Year)	Code	V	Amount	(A) or (D)	Price		6. Ownership Form: Direct (D) or Indirect (I) (Instr. 4)	7. Nature of Indirect Beneficial Ownership (Instr. 4)
Applied Biosystems Group Common Stock	11/06/2006		M		5,000	A	$ 15.54	10,578.2615	D
Applied Biosystems Group Common Stock	11/06/2006		M		12,500	A	$ 20.185	23,078.2615	D
Applied Biosystems Group Common Stock	11/06/2006		M		11,250	A	$ 21.475	34,328.2615	D
Applied Biosystems Group Common Stock	11/06/2006		S		6,250	D	$ 37.95	28,078.2615	D
Applied Biosystems Group Common Stock	11/06/2006		S		5,000	D	$ 37.85	23,078.2615	D
Applied Biosystems Group Common Stock	11/06/2006		S		12,500	D	$ 37.65	10,578.2615	D
Applied Biosystems Group Common Stock	11/06/2006		S		500	D	$ 37.54	10,078.2615	D
Applied Biosystems Group Common Stock	11/06/2006		S		700	D	$ 37.52	9,378.2615	D
Applied Biosystems Group Common Stock	11/06/2006		S		3,800	D	$ 37.5	5,578.2615	D
Applied Biosystems Group Common Stock	11/07/2006		M		1,900	A	$ 33.055	7,478.2615	D
Applied Biosystems Group Common Stock	11/07/2006		S		100	D	$ 38.45	7,378.2615	D
Applied Biosystems Group Common Stock	11/07/2006		S		200	D	$ 38.43	7,178.2615	D
Applied Biosystems Group Common Stock	11/07/2006		S		200	D	$ 38.42	6,978.2615	D
Applied Biosystems Group Common Stock	11/07/2006		S		1,400	D	$ 38.41	5,578.2615	D

Reminder: Report on a separate line for each class of securities beneficially owned directly or indirectly.
	Persons who respond to the collection of information contained in this form are not required to respond unless the form displays a currently valid OMB control number.		SEC 1474 (9-02)

Table II - Derivative Securities Acquired, Disposed of, or Beneficially Owned
(e.g., puts, calls, warrants, options, convertible securities)

1. Title of Derivative Security
(Instr. 3)

2. Conversion or Exercise Price of Derivative Security

3. Transaction Date (Month/Day/Year)

3A. Deemed Execution Date, if any (Month/Day/Year)

4. Transaction Code
(Instr. 8)

5. Number of Derivative Securities Acquired (A) or Disposed of (D)
(Instr. 3, 4, and 5)

6. Date Exercisable and Expiration Date
(Month/Day/Year)

7. Title and Amount of Underlying Securities
(Instr. 3 and 4)

8. Price of Derivative Security
(Instr. 5)

9. Number of Derivative Securities Beneficially Owned Following Reported Transaction(s)
(Instr. 4)

10. Ownership Form of Derivative Security: Direct (D) or Indirect (I)
(Instr. 4)

11. Nature of Indirect Beneficial Ownership
(Instr. 4)

Code

(A)

(D)

Date Exercisable

Expiration Date

Title

Amount or Number of Shares

Applied Biosystems Group Employee Stock Options-Right to Buy

$ 15.54

11/06/2006

5,000

03/24/2006

03/24/2013

Applied Biosystems Group Common Stock

5,000

$ 15.54

Applied Biosystems Group Employee Stock Options-Right to Buy

$ 20.185

11/06/2006

12,500

⁽¹⁾

06/17/2014

Applied Biosystems Group Common Stock

12,500

$ 20.185

12,500

Applied Biosystems Group Employee Stock Options-Right to Buy

$ 21.475

11/06/2006

11,250

06/02/2005⁽²⁾

06/02/2015

Applied Biosystems Group Common Stock

11,250

$ 21.475

33,750

Applied Biosystems Group Employee Stock Options-Right to Buy

$ 33.055

11/07/2006

1,900

⁽³⁾

11/15/2011

Applied Biosystems Group Common Stock

1,900

$ 33.055

33,100

Reporting Owners

Reporting Owner Name / Address	Relationships
Reporting Owner Name / Address	Director	10% Owner	Officer	Other
NAYYAR SANDEEP APPLERA CORP - APPLIED BIOSYSTEMS GROUP 850 LINCOLN CENTRE DRIVE FOSTER CITY, CA 94404			Assistant Controller

Signatures

/s/ Thomas P. Livingston, Attorney-In-Fact for Sandeep Nayyar		11/08/2006
^**Signature of Reporting Person		Date

Explanation of Responses:

*	If the form is filed by more than one reporting person, see Instruction 4(b)(v).
**	Intentional misstatements or omissions of facts constitute Federal Criminal Violations. See 18 U.S.C. 1001 and 15 U.S.C. 78ff(a).
(1)	These options originally became exercisable in four equal annual installments, commencing June 17, 2005. Effective January 20, 2005, the vesting of all of these options was accelerated. However, shares received upon the exercise of accelerated options are subject to a restriction on transfer (covering sales, gifts, pledges, and any other method of disposition) that remains in effect until the earlier of the original vesting dates of those options or termination of employment for any reason.
(2)	These options are fully vested and exercisable as of the grant date. However, shares acquired upon the exercise of these options are subject to a restriction on transfer (covering sales, gifts, pledges, and any other method of disposition). The transfer restriction will lapse on 25% of the shares covered by the grant on each of the first four anniversaries of the grant date. Also, the transfer restriction will lapse in full upon termination of employment for any reason.
(3)	These options originally became exercisable in four equal annual installments, commencing November 15, 2002.

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